America’s “Serengeti” at risk: Population-driven sprawl threatens the Greater Yellowstone Ecosystem

From the Everglades to the Arctic plains, population growth and associated development threaten America’s protected areas. A recent study from NumbersUSA, Greater Yellowstone: An Ecosystem at Risk, documents this for the United States’ flagship national park.

By Leon Kolankiewicz and Rob Harding

Rampant development and a surging human population are imperiling what has been dubbed “America’s Serengeti”: the iconic Greater Yellowstone Ecosystem.

Established in 1872, Yellowstone National Park was the very first in the world. While originally set aside for its unrivaled geologic and hydrothermal features (e.g., geysers and hot springs), Yellowstone has also played a critical role in the survival of such iconic wildlife as the American buffalo or bison (Bison bison), gray wolf (Canis lupus), and grizzly (brown) bear (Ursus arctos).

Bison grazing in Yellowstone National Park.
Photo by Leon Kolankiewicz

Yellowstone preserves some 2.2 million acres (about 890,000 hectares or 8,900 square kilometers) of wildlands and wilderness reaching into three states in the northern Rocky Mountains: Wyoming, Montana, and Idaho. Urban developments such as housing subdivisions, office districts, and shopping malls are all prohibited in this publicly-owned and protected national park. These habitat-destroying incursions are also banned in neighboring Grand Teton National Park, five national forests, three national wildlife refuges, and the Wind River Indian Reservation, collectively known as the Greater Yellowstone Ecosystem (GYE) or just Greater Yellowstone.

However, privately-owned lands make up about one quarter of Greater Yellowstone’s 23 million acres (93,000 square kilometers, larger than Scotland!). Most of these private lands are still rural – in a mix of cropland, pastureland, rangeland, and forest – that is, mostly cultivated farmland and ranchland used for crop and forage production and livestock (mostly cattle) grazing.

While privately-owned and often altered or degraded by agricultural practices, these typically lower-elevation lands are still ecologically crucial for the migration and over-wintering of large ungulates (herbivorous hooved mammals) such as elk or wapiti (Cervus canadensis), mule deer (Odocoileus hemionus), pronghorn (Antilocapra americana), moose (Alces alces), and bison. Even with a warming climate, the rugged mountain ranges and high elevations in the Northern Rockies still experience deep snow and harsh conditions, which many mammals have avoided since time immemorial by migrating to lower lands and habitats for the winter months.

Unfortunately, these ancient seasonal pilgrimages and survival strategies in the GYE are now threatened by a booming human population, the development and sprawl it engenders, and the attendant loss and fragmentation of crucial winter habitat. 

Squeezed out: Wintering elk in Montana’s Gallatin Valley are crowded into fragmented patches of habitat as they lose ground to sprawling residential development.
Photo by Holly Pippel

Increasing human numbers in Greater Yellowstone not only leads to permanent wildlife habitat loss, degradation, and fragmentation, but increased mortality of wildlife from collisions with vehicles on ever busier roads. This past October, arguably the most famous grizzly bear in the world, Grand Teton National Park’s Grizzly 399, a 28-year old female that was still reproductively active and known to have raised 18 cubs over her lifetime, “was fatally struck by a vehicle on Highway 26/89 in Snake River Canyon, south of Jackson, WY outside of Grand Teton National Park,” according to the National Park Service.

Already the third-most populous country on Earth in 1990 at 250 million, America’s numbers grew by 32 million in the 1990s, 27 million in the 2000s, and 22 million in the 2010s. At the peak of de facto “open borders” immigration folly under the recent Biden Administration, immigration alone added another 3.5 million in a single year (2023). If extended for a decade, and added to net natural increase (births minus deaths), this would easily exceed 40 million. Today the United States is pushing a bloated 340 million human residents and the population continues to grow with no end in sight. 

While the new Trump-Vance administration promises to slash illegal immigration, they seem supportive of increasing legal immigration, and the new vice-president has heaped scorn on “childless cat ladies.” It appears that the new administration endorses the mantra “growth is good and more is better.”

This rapid national growth has paralleled and helped propel a population boom in the Northern Rockies, as more Americans flee overpopulated, overcrowded, increasingly dysfunctional states like California for the “greener pastures” of regions that until recently, enjoyed quite low population density or what Americans call ample “elbow room”.

Grand Teton National Park Grizzly Bear 399 with one of the known 18 cubs she raised during her 28-year lifetime.
Photo by C.J. Adams, National Park Service
Dead elk killed by traffic along a busy…and ever busier…road in Greater Yellowstone.
Photo by Holly Pippel

NumbersUSA recently completed a study, called Greater Yellowstone: An Ecosystem at Risk, which quantifies the relative importance of the two main factors driving urban sprawl and the permanent conversion of rural land to developed land in the Greater Yellowstone Ecosystem. All of this rural land is wildlife habitat, agricultural land, or both.

These two main drivers of sprawl are: 1) population growth, that is, an increase in the number of residents (consumers of developed land); and 2) increase in the average consumption or use of developed land per capita, i.e., per person.  

We used a mathematical formula originally used to assess the relative weights of increasing population size and per capita energy use in determining America’s aggregate energy consumption. This approach can be extended to any natural resource, the aggregate consumption of which is increasing over time, due to a changing number of resource consumers, changing per capita resource consumption, or both. In our study, rural, undeveloped land is the natural resource in question.

Cover Photo by Holly Pippel: Bison overlooking the rapidly growing Gallatin Valley of Montana, north of Yellowstone National Park.

We used data from two federal government agencies: the Natural Resources Conservation Service (NRCS) of the U.S. Department of Agriculture and the U.S. Census Bureau (USCB). NRCS National Resources Inventories (NRIs) have estimated land use and cover on America’s non-federal lands in all 3,144 counties in the U.S. every five years since 1982. USCB estimates county populations annually. With both datasets available from 1982 to 2017 (the most recent year for the NRI), we derived estimates of the percentage of sprawl in Greater Yellowstone related to population growth and to increasing per capita developed land consumption. That is to say, if the area of sprawl increased to the same extent as population growth, with the same area of developed land per person, then the sprawl would be due solely to population growth. However, if the area of developed land per person also increases then part of the impact is due to per capita land consumption.

The area of developed non-federal land in the 20 GYE counties grew from 345,300 acres (140,000 hectares or 540 square miles) in 1982 to 497,400 acres (201,300 hectares or 777 square miles) in 2017, an increase of 44% or 152,100 acres (61,550 hectares or 237 square miles). Approximately 67% of this increase was related to population growth and 33% to increasing per capita developed land consumption. In the most recent 2002-2017 subset, 85% of the sprawl was related to population growth. These results may underestimate the adverse effects of low-density exurban sprawl on habitat fragmentation and large mammal migration. 

Sprawl Factors (Increasing Population and Increasing Per Capita Land Consumption) in GYE Counties, 1982-2017.

By 2060, the aggregate population of the GYE counties is projected to grow from approximately 540,000 in 2022 to about 765,000, an increase of about 225,000 or 42%. If average population density were to stay the same, this would lead to the loss of another 232,000 acres (94,000 hectares, or 362 square miles) of rural land (e.g., habitat, ranchland). This loss would likely entail significant adverse, long-term effects on biological diversity and abundance, in particular Greater Yellowstone’s large mammals.

Sprawl can be both low-density (left) and high-density (right); both devour habitat.
Left photo by Leon Kolankiewicz; right photo by Todd Wilkinson

As we said to the 16th Biennial Scientific Conference on the Greater Yellowstone Ecosystem in September 2024:

Avoiding this undesirable outcome will require a combination of: 1) effective local planning measures and, 2) commitment to national population stabilization. Each is necessary, neither is sufficient in itself, to preserve the unique character and iconic wildlife of the GYE.

The good news is that there is political support both among Americans generally and Greater Yellowstone residents in particular for actions to control sprawl at both the local and national levels. This was clearly revealed in the results of polling conducted last summer by Rasmussen Reports and NumbersUSA. Majorities supported both local measures such as zoning and urban growth boundaries, and national measures such as reducing the level of annual immigration admissions to slow national population growth that is the indirect or underlying driver of growth in the GYE and development pressures wildlife face. 

Both local and national measures face stiff political headwinds among elected officials, but so have many ultimately successful conservation efforts over the years. As author and veteran journalist Todd Wilkinson, founder of the conservation journalism website Yellowstonian (yellowstonian.org), wrote in the closing words to the foreword of our study:

…there is only one Yellowstone and one Greater Yellowstone Ecosystem. If we can’t succeed in protecting this bioregion, then what hope, really, do we have for saving anyplace else?

Leon Kolankiewicz is an environmental consultant, conservation biologist, vice president of Scientists and Environmentalists for Population Stabilization, and scientific director at NumbersUSA.

Rob Harding is Sustainability Outreach Liaison at NumbersUSA, board member of The Rewilding Institute, trustee of WILD Foundation, and chapter director of Center for the Advancement of the Steady State Economy.

Published

21 responses to “America’s “Serengeti” at risk: Population-driven sprawl threatens the Greater Yellowstone Ecosystem”

  1. Esther Phillips Avatar

    We have no descendants to worry about any of this.
    Their non-existence is the greatest, most loving gift we were able to give them.

  2. gaiabaracetti Avatar

    Before Yellowstone was “established”, Native Americans had lived there for thousands of years, with no need for parks, and a very thriving nature. This is true for pretty much all parks: the native people were kicked out of their lands to make room for others, and also kicked out from the supposedly “wild” areas to protect the little that was left after those newcomers had ruined everything else.
    It’s so annoying to see this crucial fact never mentioned in these articles. It’s the same story anywhere in the world. It’s not pristine nature vs humans, it’s one way of living in nature vs another.
    TOP, please keep up and update this old conservationist view. More and more research is showing how indigenous management is very often better for biodiversity than even natural parks and protected areas. We need new approaches. Until humans disappear completely, we need to look at indigenous strategies for coexistence, because we still need to eat and take things from nature in a sustainable way.

    1. gaiabaracetti Avatar

      Parks are a colonial creation. In some areas now they might be necessary given the present situation, but different alternatives should be explored
      https://www.theguardian.com/environment/2025/jan/28/conservation-environment-africa-community-wildlife-conservancies-national-parks-sustainable-tourism-indigenous-people-aoe

    2. Leon Kolankiewicz Avatar

      Gaia makes some very good points about the crucial role that indigenous peoples can and should play in protecting and managing their homelands. I myself have worked with Native American and First Nation peoples on environmental conservation and protected area projects in Central America, the American Southwest, western Canada, and Alaska.

      But whether owned and managed by indigenous peoples or “colonizers,” the direct and indirect threats that protected areas and imperiled wildlife face from mounting human population pressures and resource demands are all too real. It is unrealistic to believe that deep-rooted spiritual and traditional connections to the land and indigenous knowledge alone are enough to ward off those threats.

      Our study on the growing threat population-driven sprawl and development pose to wildlife habitat in the Greater Yellowstone Ecosystem was not intended to address the larger, deeper issues Gaia raises, but to focus on the narrower question of what intensifying population pressures signify for this world renowned treasure. These pressures are on PRIVATE lands within the ecosystem, not the publicly-owned national parks or forest lands whose very legitimacy Gaia is calling into question.

      In any case, at Yellowstone and elsewhere on public lands such as national parks, Native Americans are appropriately playing a larger and larger role in the management of all Americans’ shared natural heritage. For example, the Introduction to Yellowstone National Park’s “State of the Park 2023” report stated: “On March 1, 2022, Yellowstone turned 150. This significant milestone was commemorated throughout the year by highlighting many of the successes within the Yellowstone ecosystem, with a special emphasis on increasing our engagement with the park’s 27 associated American Indian Tribes.”

      1. gaiabaracetti Avatar

        Thank you for this reply and sorry for not addressing the more specific points your raised and going on a little rant. I absolutely agree with population stabilisation, of course. It’s interesting, actually, that – unless I’m badly informed – advocates for indigenous rights don’t see the immigrants as a threat to the land and environment, but the mainstream capitalist industrial society. It’s hard to untangle and even a little contradictory. That’s why discussions about population and economic growth need to go hand in hand, otherwise we’re always going to hear “there could be more of us here, if only we lived with less”. Which is not going to work.

    3. Oats n groats Avatar

      Indigenous tribes only managed the land by unintentionally having low populations through high maternal and infant mortality rates and lots of attrition through war with other tribes. You’re propagating the old myth of the “noble savage”. If natives had had modern medicine, their numbers would have swelled and they would have destroyed the environment just as successfully as whitey does. People are people.

      1. gaiabaracetti Avatar

        No, this is not entirely true. Many human societies had developed various forms of birth control before modern medicine. Also, the numbers of natives in places that were then colonised were much higher than today. The Amazon had thousands of cities: https://www.theguardian.com/global-development/2025/feb/06/ancient-garden-cities-amazon-indigenous-technologies-archaeology-lost-civilisations-environment-terra-preta This doesn’t seem to have destroyed the environment.
        Mass extinctions are another matter, but those go way back and happened everywhere except for Africa.
        As for managing the land, there’s evidence that through things like controlled burns, cultivation and domestication, hunting, fishing, etc, many native peoples “managed” their environments in ways that could be destructive, but also had the effect of increasing biological productivity or biodiversity.
        Of course people are people, but some systems are more sustainable than others. The non-sustainable ones, in any continent, collapsed.

  3. PHILIP CAFARO Avatar

    National Parks, wilderness areas and other kinds of protected areas are not “colonial creations,” they are a response to modern realities. If we want to preserve other species, we need to leave them the habitat and resources — the ecological space — to do so. If we want to have some landscapes that are not primarily human creations, we have to set them aside.

    The alternative to national parks and protected areas is not a return to primitive or premodern forms of life, it is allowing industrial capitalism to run rampant over 100% of the landscape. While indigenous control of the landscape can sometimes lead to less heavy handed use, it can also facilitate rape and pillage by energy and mining companies, when indigenous groups cut deals with them to sacrifice their ancestral lands for profit.

    1. gaiabaracetti Avatar

      In most areas of the world, though, those modern realities were created precisely by colonisation. I think this factor should not be forgotten or overlooked. Before colonialism (not just by white people of course), all those protected areas had people living in them, and in most cases nature was thriving. It wasn’t all ideal, sure, but it was much better than now. Protection was very localised, specific sacred or off-limits places, but you didn’t need to set aside large swathes of land like we do now with parks.
      As for energy and mining conflicts, do you have specific examples in mind of indigenous people collectively sacrificing their ancestral lands for profit? In every single case that I can think of or have read about, in the Americas, Asia, Africa, Northern Europe, it’s the indigenous people fighting for protection of the land, rather than viceversa. The people who want to “develop” or plunder the land usually come from outside. I can think of some exceptions for example in some Pacific islands that are desperately poor, but even there, they have marine reserves that are locally managed.
      It’s not just a cultural thing; it’s the fact that you know that if you lose the land, you lose your livelihood and home, whereas industrial capitalism is stateless and can just move somewhere else once it’s wrecked a place. Migrants, also, move to the Amazon or the US or Northern Europe because they need a place to go, but it doesn’t have to be *that* specific place, it could just be anywhere as long as it meets their criteria.
      And even here, for example in the Alps, the pillaging often goes hand in hand with the displacement of native populations. Hydro is one example. Logging is another. Locals traditionally used water and cut wood much more responsibly, knowing that they had to last. In the places where they didn’t, the land was abandoned eventually because it could no longer sustain the population. There’s a self-selection of sorts in place that is quite “natural”. That’s why indigenous people usually have such precious knowledge.

      1. PHILIP CAFARO Avatar

        Unfortunately, there are lots of examples of native peoples in North America sacrificing their lands for profit by leasing them to mining and energy companies, lots of examples of tribes opposing the reintroduction of native species such as wolves, lots of examples of them opposing protected area designation because they want to exploit lands rather than protect them.

        Here are a few:

        https://www.nytimes.com/2024/12/13/climate/canada-lng-natural-gas-indigenous-first-nations.html

        https://coloradosun.com/2024/08/06/wolves-colorado-washington-tribe/

        https://www.montanarightnow.com/news/navajo-transitional-energy-company-takes-over-mines-in-montana-wyoming/article_b98768f0-da79-5a61-a164-9bab525743cd.html

        That said, there are also great examples of native american land stewardship. Indians are people, too, and do good and bad things. No surprise there!

  4. Jane O'Sullivan Avatar

    In many countries, indigenous people are actively involved in managing, and advising on the management of, national parks. This generally doesn’t mean that they live off the landscape and harvest it in the way their ancestors did. They wouldn’t choose that lifestyle now even if it were viable, but usually it is not viable because the areas being conserved are not big or diverse enough. I think it is unfair to blame “colonialism” when often endemic population growth is to blame for most of the encroachment on natural environments. It is also unhelpful: we can’t undo history. The descendants of colonists can be just as committed to protecting wild places as indigenous people.

    1. PHILIP CAFARO Avatar

      I would add that it is up to all of us to “become indigenous” to our own places and nations now. We need to learn to know, appreciate and protect the places where we live.

      Who’s to say Robin Kimmerer, the author of “Braiding Sweetgrass,” who grew up in a conventional American town thinking in and speaking English, but has some amount of native American genes and registered with an Indian tribe as an adult, is more indigenous to North America than me? Or that Gaia isn’t more indigenous (to northern Italy, in her case) than either of us, spending more time working the land?

      1. gaiabaracetti Avatar

        Of course, and I’m not a big fan of treating white Americans (or Canadians, Australians, etc) as perennial colonisers. After a while, we’re all native; if we go far back enough, none of us are. Also, belonging is earned, not just given; some people have no loyalty to the place they were born, and others would give their life for a place where they were not.
        What I was talking about was more the recognition of this fact and an ending of the erasure of the history and human presence on those specific lands. Also, in places like US, Australia, Latin America, the descendants of colonisers and the richer immigrants own unbelievably large swathes of land that belongs exclusively to them in a way that looks absurd to both an European and an indigenous person from those lands. When I hear about properties being hundreds or thousands of acres I think – what? How is this even legal?
        It’s interesting how no one thinks of white Europeans in Europe as “indigenous”, even though that’s exactly what most of us are. Even more interesting is to study the conflict between the aristocratic elites and the lower classes that often do resemble those between “colonisers” and natives. For example, I’ve learnt that here people would destroy “developments” carried out by rich absentee landlords, and protect things like swamps, communal pastures, fishing grounds… it took centuries to break them, with a carrot-and-stick approach. Scottish people kicked off the land and put on boats to Canada to make room for sheep monocultures are a similar story.
        Finally, this indigenous-non indigenous dichotomy also applies, like I was saying, to the indigenous people of India (the Adivasi), many non-Han Chinese, and many others. It’s not a race-based division but an economic system and expansionist mindset division, which is what I was trying to get at.

  5. gaiabaracetti Avatar

    Philip, that’s still not an argument for stripping people of sovereignty, which is an inalienable right. The same way we want to have our democracy and right to vote even if we make bad choices. I can’t read any of the articles because they are either not available to me or behind paywalls, so I can’t comment on those specifically. When I was living in Canada I learned how the First Nations were displaced against their will for massive hydro projects that produced a lot of energy and completely wrecked the environment and their ability to take sustenance from it; in Western Canada they are fighting against industrial logging and trying to protect the forests. I don’t know about the gas, I’ll look it up.
    The opposition to the protected area designation is often justified because people want to be able to live off their land sustainably, and protection might prevent that. You are framing this in terms of “exploitation” but people do need to eat, warm themselves, make the things they need, in a culturally appropriate way. If you prevent them completely from doing this on their own land, they will become dependent on an economy they don’t want to participate in and on an alien culture. You criticise them but you are secure in your livelihood, which allows you to purchase things, while other people want to make them themselves or sell them in order to support their families. If you are asking them to give this up, what are you giving in return? So it’s not the principle of not wanting full protection that is wrong, but the choices you make about the limits that you do accept.
    Being opposed to the reintroduction of wolves does not count in my book as “sacrificing the land for profit”. I think that reintroductions are often experiments made on other people’s lands in a way that threatens their livelihood without offering anything in return. A balance should be restored anywhere that it’s possible, cities included, and then species should come back on their own if the environment is suitable, which is what happens for example in some areas in Europe. Buffaloes are an example of a Native-managed reintroduction that is restoring the prairie ecosystem, but buffaloes they can kill and eat, unlike wolves. In other areas, anyway, native people protect predators from killing, so it’s a mixed bag and depends on the context.
    This is the colonial mentality I was talking about: people who’ve been deprived of their traditional livelihood by a majority population that has absolutely wrecked the environment thus acquired, and who are trying to hold on to some land and autonomy, being constantly judged and considered unworthy by that same society that has done the major damage. Would you support putting wolves in Central Park? Why not? Why should some areas be kept prosperous and safe, while others, in which other people live and try to make a living too, be subjected to your idea of what the world should look like? Start with your own city, space, job, then tell others what to do with theirs.
    I don’t disagree that indigenous people can make bad choices, like all, of course. I’m just saying that if we show ourselves willing to take a loss in exchange for asking others to do the same, or willing to redress an ancient wrong, it’s easier to be persuasive.

    1. gaiabaracetti Avatar

      P.S. It’s not a “you” as in Philip, it’s a general “you” 🙂

    2. PHILIP CAFARO Avatar

      Well, Indians have been in North America for 14 or 15 thousand years, wolves for three quarters of a million years. So I think they have a right to be here, if anyone does. Whether or not it’s convenient for people. And lots of Native Americans agree with me!

      1. gaiabaracetti Avatar

        Again, I don’t think you are making a convincing argument for depriving people of sovereignty over their own land.
        I don’t think anyone is saying wolves don’t have a “right” to exist as such.
        You either believe that deliberately causing extinction is wrong, or you believe that it’s right, in which case it doesn’t matter how long a species has been around. I personally believe deliberately causing extinction is wrong for anything, including mosquitoes.
        But reintroduction is not the same as protection, as allowing a species to exist: it’s humans deciding exactly where it should be, irrespective of the wishes of the species itself and of the new ecological equilibrium that has formed in the meanwhile. It’s a heavy-handed ecological intervention that follows human preferences. Once a predator disappears, some other animal (including but not limited to humans) fills the niche; if the persecuted predator is capable of filling that niche again, it comes back on its own once the environment allows it. The biological history of Earth is full of animals evolving to fill new niches after others have left for whatever reason. Reintroduction is humans saying: we don’t like how nature is taking care of this, or how long it’s taking to find a new equilibrium – we want that specific species in that specific place right now. It often backfires.
        So: why aren’t we reintroducing wolves to New York? Why aren’t we reintroducing them on Long Island or in other places where rich people have holiday homes and therefore can afford a loss more than people who depend on the containment of predators for survival?
        “Whether or not it’s convenient for people” only seems to apply to certain kinds of people. And I think that these kind of people, whose convenience, unlike your own, you deem irrelevant, have every right to resist.

  6. gaiabaracetti Avatar

    Sorry, just to clarify, there’s two separate issues here: sovereignty and reintroductions. Refusing reintroductions doesn’t invalidate sovereignty, which was what you implied; if a sovereign people wants to do reintroduction, it’s their choice regardless of what Gaia thinks.

    1. PHILIP CAFARO Avatar

      This could only have been written by someone with no concern for the well-being of other species, or the responsibility of American citizens to protect the wildlife we share the landscape with.

      1. gaiabaracetti Avatar

        This is unfair. Did you read the article until the end?

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